Input vs. output preferred orientation of neurons in the network. (A-C) Output PO vs. Input PO for three values of recurrence, at the medium contrast. Increasing the recurrence leads to more scatter about the diagonal. For illustration purposes, the excitatory (red) and inhibitory (blue) neurons have been plotted only above or below the diagonal, respectively. (D) To quantify the amount of PO change when going from input to output, the Scatter Degree Index (SDI) is plotted as the angular deviation of Δ PO = OutputPO−InputPO (see Methods). The maximum value of this index is ≈40.5°, which corresponds to a uniform distribution of Δ PO. Darker colors show higher contrasts, respectively.