An omnibus permutation test on ensembles of two-locus analyses can detect pure epistasis and genetic heterogeneity in genome-wide association studies

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Table 6 Two-locus penetrances that lead to the heritability of 0.01

AA and BB denote homozygous wild-type genotypes. Aa and Bb denote heterozygous genotypes. aa and bb denote homozygous variant genotypes. All allele frequencies are equal (p _A = p _B = 0.5). K = 1/201.